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Biology Year 12 Module 5 Lesson 15

Non-Mendelian Patterns - Co-dominance, Incomplete Dominance, Multiple Alleles

Not every inheritance pattern fits a simple dominant-recessive model. Some traits show both alleles together, some show an intermediate phenotype, and some genes have more than two alleles in the population.

40 min IQ4 Genetic variation 5 MC | 3 Short Answer Lesson 15 of 19
ABO
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Prediction

Think First

A student says, "If red flower colour is dominant and white flower colour is recessive, then crossing a red flower with a white flower should always give red offspring. So pink flowers prove the experiment went wrong."

Before reading on, explain why that reasoning is weak. What inheritance pattern could produce pink offspring, and why does that not mean the alleles have disappeared or blended permanently?

Key Terms
Co-dominanceBoth alleles are fully expressed in the heterozygous phenotype.
Incomplete dominanceThe heterozygous phenotype is intermediate between the two homozygous phenotypes.
Multiple allelesA gene exists in more than two allele forms within the population.
HeterozygousHaving two different alleles for a gene.
ABO blood groupA human blood group system showing co-dominance and multiple alleles.
Phenotypic ratioThe relative frequency of observable traits in offspring.

Know

  • What co-dominance, incomplete dominance and multiple alleles mean.
  • That ABO blood groups are the standard HSC example of co-dominance plus multiple alleles.

Understand

  • Why some phenotype ratios are not the classic 3:1 Mendelian ratio.
  • Why intermediate phenotype does not mean the alleles have merged permanently.

Be Able To

  • Interpret Punnett squares for non-Mendelian patterns.
  • Distinguish co-dominance from incomplete dominance in examples and data.

Misconceptions to Fix

Wrong: Homeostasis means the body stays exactly the same all the time.

Right: Homeostasis involves dynamic equilibrium — constant small adjustments around a set point.

1
Beyond Simple Dominance

Not all heterozygotes look like one homozygote

Simple dominant-recessive inheritance is only one inheritance pattern. The effect of two alleles in a heterozygote depends on how the gene product is expressed.

In Lesson 14, the heterozygous genotype often showed the dominant phenotype. In this lesson, the heterozygous phenotype may be intermediate, or it may show both allele products together. That is why some crosses do not produce the expected 3:1 phenotypic ratio.

Simple Mendelian

One allele masks the other in the heterozygote, often producing a 3:1 phenotype ratio.

Incomplete dominance

The heterozygote shows an intermediate phenotype, often producing a 1:2:1 phenotype ratio.

Co-dominance

Both alleles are expressed in the heterozygote, so the phenotype includes both.

Trap
Do not confuse co-dominance with incomplete dominance. Co-dominance means both allele products are visible together. Incomplete dominance means the heterozygous phenotype is intermediate.
2
Intermediate Phenotype

Incomplete dominance gives a blended phenotype, not blended inheritance

In incomplete dominance, neither allele is fully dominant in the heterozygote. A common example is red-flowered and white-flowered snapdragons producing pink heterozygous offspring.

CR CW CR CW CRCR CRCW CRCW CWCW Cross: CRCW x CRCW Genotypes: 1 red : 2 pink : 1 white Phenotypes: 1 red : 2 pink : 1 white
In incomplete dominance, the 1:2:1 genotype ratio often matches the 1:2:1 phenotype ratio.

The key idea is that the alleles still segregate normally in meiosis. They have not permanently mixed into one new allele. The intermediate phenotype appears because of the expression pattern in the heterozygote.

3
Both Alleles Visible

Co-dominance means both allele products are expressed

In co-dominance, the heterozygous phenotype shows both allele products. Neither allele hides the other, and the phenotype is not simply halfway between them.

Co-dominance

  • Both alleles are expressed in the heterozygote.
  • The phenotype shows both forms together.
  • AB blood type is the standard human example.

Incomplete dominance

  • The heterozygote has an intermediate phenotype.
  • It does not show both separate products clearly together.
  • Pink flowers are the standard school example.
Example
In the ABO blood group, the IA and IB alleles are co-dominant. A person with genotype IAIB has blood group AB because both A and B antigens are expressed on the red blood cells.
4
Population View

Multiple alleles means more than two allele forms exist in the population

Each individual still has only two alleles for an autosomal gene, one inherited from each parent. However, the gene may exist in more than two forms across the population. This is called multiple alleles.

ABO alleles

The ABO blood group has three common alleles: IA, IB and i.

Dominance relationships

IA and IB are co-dominant with each other, and both are dominant over i.

Possible blood groups

Phenotypes are A, B, AB and O, depending on which two alleles the individual has.

Phenotype A Phenotype B Phenotype AB Phenotype O IAIA IAi IBIB / IBi IAIB ii
ABO shows both multiple alleles in the population and co-dominance between IA and IB.
5
Interpretation

Observed ratios tell you which inheritance pattern is likely

If the heterozygote shows the dominant phenotype, a monohybrid cross often gives a 3:1 phenotype ratio. If the heterozygote is intermediate, the same cross may give a 1:2:1 phenotype ratio. That is why phenotype data matters in inheritance questions.

When you see 3:1

  • Think simple dominant-recessive inheritance.
  • The heterozygote likely matches one homozygote phenotypically.

When you see 1:2:1

  • Consider incomplete dominance or co-dominance.
  • The heterozygote likely has its own distinct phenotype.
Exam Rule
Do not assume every monohybrid cross gives a 3:1 phenotype ratio. First identify how the heterozygote is expressed, then interpret the Punnett square.
Copy Into Your Books +

Co-dominance

In co-dominance, both alleles are fully expressed in the heterozygous phenotype, such as IAIB giving blood group AB.

Incomplete dominance

In incomplete dominance, the heterozygous phenotype is intermediate between the two homozygous phenotypes, such as red and white flowers producing pink offspring.

Multiple alleles

Multiple alleles means more than two allele forms exist in the population, even though each individual still has only two alleles for the gene.

Ratios

Non-Mendelian inheritance patterns often produce phenotype ratios other than 3:1 because the heterozygote has its own distinct phenotype.

Revisit Your Initial Thinking

Look back at what you wrote in the Think First section. What has changed? What did you get right? What surprised you?

Activities

Activity 1: Flower colour cross

In a plant species, red flowers and white flowers show incomplete dominance. Heterozygous plants are pink.

Cross two pink plants. Write the Punnett square, then state the genotype and phenotype ratios.

Activity 2: ABO reasoning

One parent has genotype IAi and the other has genotype IBi.

List all possible offspring genotypes and identify the possible blood group phenotypes. Explain why this example shows both multiple alleles and co-dominance.

Multiple Choice

Understand 1 mark

1. What best describes co-dominance?

A
One allele completely masks the other
B
Both alleles are fully expressed in the heterozygote
C
The heterozygote is always identical to one parent
D
More than two chromosomes carry the gene
Understand 1 mark

2. Which statement about incomplete dominance is correct?

A
The heterozygous phenotype is intermediate between the two homozygous phenotypes
B
Both alleles disappear in the heterozygote
C
It is another name for sex linkage
D
It only occurs in plants and never in animals
Apply 1 mark

3. A gene has three common allele forms in the population. What term describes this?

A
Incomplete dominance
B
Co-dominance
C
Multiple alleles
D
Polygenic inheritance
Analyse 1 mark

4. Which genotype gives blood group AB in the ABO system?

A
ii
B
IAi
C
IAIB
D
IBi
Analyse 1 mark

5. A cross between two heterozygotes produces a 1:2:1 phenotype ratio. What does this most strongly suggest?

A
The trait is definitely X-linked
B
The heterozygote has a distinct phenotype, such as in incomplete dominance or co-dominance
C
The alleles have fused permanently into one new allele
D
One parent must be homozygous recessive

Short Answer

Apply 3 marks

6. Explain the difference between co-dominance and incomplete dominance using one example for each.

3 marks

Apply 4 marks

7. Two pink flowers are crossed in a species showing incomplete dominance. Red flowers are CRCR, white flowers are CWCW, and pink flowers are CRCW.

4 marks

Determine the genotype ratio and phenotype ratio of the offspring.

Analyse 5 marks

8. In the ABO blood group system, one parent has genotype IAi and the other has genotype IBi.

5 marks

Use Punnett square reasoning to identify all possible offspring genotypes and phenotypes, and explain why this system demonstrates both co-dominance and multiple alleles.

Rapid Review

Co-dominance

The heterozygote expresses both allele products, as in blood group AB.

Incomplete dominance

The heterozygote has an intermediate phenotype, as in pink flowers from red and white parents.

Multiple alleles

A gene can have more than two allele forms in the population even though each individual still carries only two alleles.

Revisit Your Thinking

Return to the pink-flower claim from the start of the lesson. Rewrite it using the correct inheritance language.

Answers and Worked Solutions

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Multiple Choice

1. B - Co-dominance means both alleles are expressed in the heterozygote.

2. A - In incomplete dominance, the heterozygote is intermediate between the two homozygotes.

3. C - More than two allele forms in the population is called multiple alleles.

4. C - Blood group AB has genotype IAIB.

5. B - A 1:2:1 phenotype ratio suggests the heterozygote has its own distinct phenotype.

Short Answer 6

Co-dominance means both alleles are fully expressed in the heterozygote, such as IAIB producing blood group AB. Incomplete dominance means the heterozygote shows an intermediate phenotype, such as pink flowers from a red and white flower cross.

Short Answer 7

The cross is CRCW x CRCW. Each parent produces gametes CR and CW. Offspring genotypes are CRCR, CRCW, CRCW and CWCW. Genotype ratio = 1 CRCR : 2 CRCW : 1 CWCW. Phenotype ratio = 1 red : 2 pink : 1 white.

Short Answer 8

The first parent produces gametes IA and i. The second parent produces gametes IB and i. Possible offspring genotypes are IAIB, IAi, IBi and ii. Their phenotypes are AB, A, B and O. This shows co-dominance because IA and IB are both expressed together in blood group AB. It shows multiple alleles because the ABO gene exists as three common alleles in the population: IA, IB and i.

Mark lesson complete

Tick this once you can distinguish co-dominance, incomplete dominance and multiple alleles without forcing every question into a simple 3:1 pattern.