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hscscience Ext 1 · Y12
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Module 9 · L15 of 20 ~40 min ⚡ +95 XP available

The Logistic Equation

A petri dish starts with 100 bacteria. They double every hour, but the dish can only hold 10 000. Does growth just stop at capacity? What does the population curve look like? The logistic equation $\dfrac{dP}{dt} = kP\!\left(1 - \dfrac{P}{M}\right)$ captures this S-shaped reality. In this lesson you'll solve it using partial fractions, sketch the solution curve, and answer HSC-style questions on carrying capacity and inflection.

Today's hook, A population starts at $P_0 = 200$ in an environment with carrying capacity $M = 1000$. Growth rate $k = 0.4$. Will the population ever exceed 1000? At roughly what time does growth feel fastest? Sketch a rough curve before reading card 05.
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Recall, your gut answer first
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A population starts at $P_0 = 200$ with carrying capacity $M = 1000$ and growth rate $k = 0.4$. Without using a formulasketch or describe what you expect the population-vs-time curve to look like. Does growth speed up, slow down, or both?

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The big idea
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Exponential growth ($\tfrac{dP}{dt} = kP$) predicts unlimited growth, unrealistic for any real population. The logistic model adds a braking term $\bigl(1 - \tfrac{P}{M}\bigr)$ that shrinks to zero as $P \to M$:

When $P \ll M$ the factor $\approx 1$ and growth is nearly exponential. When $P \approx M$ the factor $\approx 0$ and growth stalls. The inflection point fastest growth, occurs at exactly $P = M/2$.

$\dfrac{dP}{dt} = kP\!\left(1 - \dfrac{P}{M}\right)$

t P M M/2 inflection
$\dfrac{dP}{dt} = kP\!\left(1 - \dfrac{P}{M}\right)$
Carrying capacity $M$
$M$ is the maximum sustainable population. As $P \to M$ from below, $\tfrac{dP}{dt} \to 0$, growth asymptotically approaches (but never exceeds) $M$.
Inflection at $P = M/2$
The rate of change of $P$ is maximised when $P = M/2$. This is the inflection point of the S-curve, growth is fastest here, then decelerates.
Solve via partial fractions
Separating variables gives $\displaystyle\int \frac{dP}{P(1-P/M)} = \int k\,dt$. The left side requires partial fractions to integrate.
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What you'll master
Know

Key facts

  • Logistic DE: $\tfrac{dP}{dt} = kP(1 - P/M)$; $M$ is carrying capacity
  • Inflection point at $P = M/2$; solution is an S-curve
  • General solution: $P(t) = \dfrac{M}{1 + Ae^{-kt}}$ where $A = \dfrac{M - P_0}{P_0}$
Understand

Concepts

  • Why the braking factor $(1 - P/M)$ produces bounded growth
  • How partial fractions decompose $\tfrac{1}{P(M-P)}$
  • The biological meaning of $k$, $M$, and the inflection point
Can do

Skills

  • Solve the logistic DE by separation of variables and partial fractions
  • Apply initial conditions to find $A$ and interpret the solution
  • Identify $k$, $M$, $P_0$ from a context and find $P(t)$
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Key terms
Logistic equationA differential equation $\tfrac{dP}{dt} = kP(1-P/M)$ modelling population growth bounded by a carrying capacity.
Carrying capacity $M$The maximum sustainable population size. The solution $P(t)$ approaches $M$ as $t \to \infty$.
Intrinsic growth rate $k$The proportional growth rate when population is far below capacity ($P \ll M$). Units: per unit time.
Inflection pointThe point on the S-curve where growth rate is maximum, occurring at $P = M/2$.
Partial fractionsA technique to decompose $\tfrac{1}{P(M-P)}$ into $\tfrac{1}{M}\!\left(\tfrac{1}{P} + \tfrac{1}{M-P}\right)$ to enable integration.
Sigmoid / S-curveThe characteristic shape of the logistic solution: slow start, rapid middle growth, levelling off near $M$.
05
Solving the logistic equation
core concept

Start with $\dfrac{dP}{dt} = kP\!\left(1 - \dfrac{P}{M}\right)$. Rewrite as $\dfrac{dP}{dt} = \dfrac{k}{M} P(M - P)$ and separate variables:

$$\int \frac{dP}{P(M-P)} = \int \frac{k}{M}\,dt$$

Partial fractions: Write $\dfrac{1}{P(M-P)} = \dfrac{A}{P} + \dfrac{B}{M-P}$. Multiplying both sides by $P(M-P)$ and comparing coefficients gives $A = \tfrac{1}{M}$, $B = \tfrac{1}{M}$. So:

$$\frac{1}{M}\ln P - \frac{1}{M}\ln(M-P) = \frac{k}{M}t + C$$

Multiplying both sides by $M$ and exponentiating:

$$\frac{P}{M - P} = Ae^{kt} \quad \Rightarrow \quad \boxed{P(t) = \frac{M}{1 + Ae^{-kt}}}$$

where $A = \dfrac{M - P_0}{P_0}$ is determined by the initial condition $P(0) = P_0$.

Check the long-run behaviour. As $t \to \infty$, $e^{-kt} \to 0$, so $P(t) \to \dfrac{M}{1+0} = M$. The population approaches carrying capacity but never exceeds it, exactly as predicted by the S-curve shape.

Start with $\dfrac{dP}{dt} = kP\!\left(1 - \dfrac{P}{M}\right)$. Rewrite as $\dfrac{dP}{dt} = \dfrac{k}{M} P(M - P)$ and separate variables:

Pause, copy the logistic solution $P(t)=\frac{M}{1+Ae^{-kt}}$ with $A=(M-P_0)/P_0$, and the partial-fraction step that generates it into your book.

Quick check: A population satisfies $\dfrac{dP}{dt} = 0.3P\!\left(1 - \dfrac{P}{500}\right)$ with $P(0) = 50$. What is the value of $A$ in the general solution $P(t) = \dfrac{M}{1+Ae^{-kt}}$?

06
The inflection point and maximum growth rate
core concept

We just saw that separating $\frac{dP}{dt}=kP(1-P/M)$ and using partial fractions gives $P(t)=\frac{M}{1+Ae^{-kt}}$ where $A=(M-P_0)/P_0$. That raises a question: the growth rate $\frac{dP}{dt}$ starts small, rises, then falls back to zero, at what exact value of $P$ is it maximised, and what is that maximum rate? This card answers it → the inflection occurs at $P=M/2$, and the maximum growth rate is $\frac{dP}{dt}=kM/4$.

The inflection point of $P(t)$ is where $\dfrac{d^2P}{dt^2} = 0$, i.e.\ where the growth rate $\dfrac{dP}{dt}$ is a maximum.

Define $f(P) = kP(1-P/M)$. Differentiate with respect to $P$:

$f'(P) = k - \dfrac{2kP}{M} = 0 \quad \Rightarrow \quad P = \dfrac{M}{2}$

At this population the growth rate equals $\dfrac{dP}{dt}\Big|_{P=M/2} = k \cdot \dfrac{M}{2} \cdot \dfrac{1}{2} = \dfrac{kM}{4}$.

HSC exam tip. Examiners love asking "at what population is the growth rate greatest?" The answer is always $P = M/2$. If they ask for the maximum rate, substitute into $\tfrac{dP}{dt}$ to get $kM/4$.

The inflection point of $P(t)$ is where $\dfrac{d^2P}{dt^2} = 0$, i.e.\ where the growth rate $\dfrac{dP}{dt}$ is a maximum.

Pause, copy the inflection-point result: $P=M/2$ gives maximum growth rate $kM/4$; above $M/2$ the growth decelerates; $P=M$ is the asymptote into your book.

Did you get this? True or false: for the logistic equation with $M = 800$, the population growth rate is greatest when $P = 400$.

PROBLEM 1 · FIND THE SOLUTION

A fish population satisfies $\dfrac{dP}{dt} = 0.2P\!\left(1 - \dfrac{P}{1000}\right)$ with $P(0) = 100$. Find $P(t)$.

1
$M = 1000$, $k = 0.2$, $P_0 = 100$. Use $P(t) = \dfrac{M}{1 + Ae^{-kt}}$.
Identify the parameters from the differential equation and initial condition.
PROBLEM 2 · INFLECTION TIME

Using the solution from Problem 1, at what time $t$ does the population reach the inflection point?

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Inflection occurs at $P = M/2 = 500$. Set $\dfrac{1000}{1 + 9e^{-0.2t}} = 500$.
The inflection point is at $P = M/2$ for the logistic equation.
PROBLEM 3 · SHOW THAT (2-PART)

Show that $P(t) = \dfrac{M}{1 + Ae^{-kt}}$ satisfies $\dfrac{dP}{dt} = kP\!\left(1 - \dfrac{P}{M}\right)$.

1
Let $Q = 1 + Ae^{-kt}$, so $P = M/Q$. Then $\dfrac{dP}{dt} = -\dfrac{M}{Q^2}\cdot(-kAe^{-kt}) = \dfrac{MkAe^{-kt}}{Q^2}$.
Differentiate using the chain rule (quotient rule with constant numerator).

Fill the gap: For the logistic equation with $M = 600$ and $k = 0.5$, the maximum growth rate equals $\dfrac{kM}{4} = $ .

Trap 01
Forgetting to include $P_0$ to find $A$
The constant $A$ is not arbitrary, it is fixed by the initial condition. Always substitute $t = 0$, $P = P_0$ into $P(t) = \tfrac{M}{1+Ae^{-kt}}$ and solve: $A = \tfrac{M - P_0}{P_0}$. Missing this step gives an incomplete answer.
Trap 02
Wrong inflection point
The inflection is at $P = M/2$, not $t = M/2$. Examiners write "at what population value is growth fastest?", the answer is $M/2$. Substituting back to find $t$ requires solving $P(t) = M/2$, giving $t = \tfrac{\ln A}{k}$.
Trap 03
Sign errors in partial fractions
$\displaystyle\int \frac{dP}{M-P} = -\ln|M-P| + C$ (negative sign because the denominator decreases). Dropping this sign flips the solution. Always differentiate your answer to verify it satisfies the original DE.

Did you get this? True or false: for a logistic population with $P_0 = M/2$, the constant $A$ in $P(t) = M/(1+Ae^{-kt})$ equals 1.

Work mode · how are you completing this lesson?
1

A population satisfies $\tfrac{dP}{dt} = 0.1P(1 - P/2000)$ with $P(0) = 200$. Write down the values of $k$, $M$, and $A$, then state $P(t)$.

2

For the DE in Activity 1, at what population value is the growth rate a maximum? What is that maximum growth rate?

3

Use partial fractions to show that $\displaystyle\int \frac{dP}{P(1000-P)} = \frac{1}{1000}\ln\left|\frac{P}{1000-P}\right| + C$.

4

A population with $M = 500$, $k = 0.6$, $P(0) = 50$ satisfies the logistic model. Find the time when $P = 400$. Give your answer in exact form.

5

Explain why a logistic population with $P_0 > M$ (population starts above carrying capacity) would decrease towards $M$. What is the sign of $\tfrac{dP}{dt}$ in this case?

Odd one out: Three of these statements about the logistic equation are correct. Which one is NOT?

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Revisit your thinking

Earlier you sketched or described the population curve for $P_0 = 200$, $M = 1000$, $k = 0.4$.

The exact solution is $P(t) = \dfrac{1000}{1 + 4e^{-0.4t}}$ (here $A = \tfrac{1000-200}{200} = 4$). Growth accelerates up to $P = 500$ (inflection at $t = \tfrac{\ln 4}{0.4} \approx 3.47$), then decelerates, asymptoting to 1000. Did your sketch capture the S-shape with deceleration near $M$?

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Multiple choice
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Short answer
ApplyBand 32 marks

Q1. Write down the general solution of $\dfrac{dP}{dt} = 0.4P\!\left(1 - \dfrac{P}{800}\right)$ with $P(0) = 160$. (2 marks)

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ApplyBand 43 marks

Q2. For the logistic equation in Q1, find the time at which the growth rate is a maximum. Give your answer in exact form. (3 marks)

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AnalyseBand 53 marks

Q3. Show that if $P(t) = \dfrac{M}{1+Ae^{-kt}}$, then $\dfrac{dP}{dt} = \dfrac{k}{M}P(M-P)$. (3 marks)

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Comprehensive answers (click to reveal)

Activity answers: 1. $k=0.1$, $M=2000$, $A=(2000-200)/200=9$, $P(t)=2000/(1+9e^{-0.1t})$  ·  2. Inflection at $P=1000$; max rate $=0.1\times2000/4=50$ per unit time  ·  3. Partial fractions: $A=B=1/1000$; integrate to get $\tfrac{1}{1000}\ln|P/(1000-P)|+C$  ·  4. $A=(500-50)/50=9$; set $400=500/(1+9e^{-0.6t})$, solve $e^{-0.6t}=1/36$, $t=\tfrac{\ln 36}{0.6}=\tfrac{5\ln 36}{3}$  ·  5. When $P>M$, $(1-P/M)<0$ so $dP/dt<0$, population decreases towards $M$.

Q1 (2 marks): $A=(800-160)/160=4$ [1]. $P(t)=\dfrac{800}{1+4e^{-0.4t}}$ [1].

Q2 (3 marks): Inflection at $P=M/2=400$ [1]. Set $\dfrac{800}{1+4e^{-0.4t}}=400$: $1+4e^{-0.4t}=2$, $e^{-0.4t}=\tfrac{1}{4}$ [1]. $t=\dfrac{\ln 4}{0.4}=\dfrac{5\ln 4}{2}$ [1].

Q3 (3 marks): Differentiate $P=M/(1+Ae^{-kt})$: $dP/dt = MkAe^{-kt}/(1+Ae^{-kt})^2$ [1]. Note $P = M/(1+Ae^{-kt})$, so $P/M = 1/(1+Ae^{-kt})$, hence $1-P/M = Ae^{-kt}/(1+Ae^{-kt})$ [1]. Multiply: $\tfrac{k}{M}P(M-P) = k \cdot \tfrac{M}{1+Ae^{-kt}} \cdot \tfrac{MAe^{-kt}}{1+Ae^{-kt}} \cdot \tfrac{1}{M} = \tfrac{MkAe^{-kt}}{(1+Ae^{-kt})^2} = dP/dt$ ✓ [1].

01
Boss battle · The Logistic Master
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Five timed questions. Beat the boss to bank a tier, gold (90% + speed), silver (75%), or bronze (50%). Replays welcome.

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02
Science Jump · platform challenge

Climb platforms by answering logistic equation questions. Lighter alternative to the boss.

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